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BioRock ISS Basalt Biomining Consortium

A defined three-species bacterial panel flown as the ESA BioRock experiment on board the International Space Station to test microbial bioleaching of economically important elements from basaltic rock under microgravity, simulated Mars gravity, and simulated Earth gravity (with ground 1 x g controls). Sphingomonas desiccabilis CP1D (DSM 16792), Bacillus subtilis NCIB 3610 (DSM 10), and Cupriavidus metallidurans CH34 (DSM 2839) were each cultivated on slides of olivine basalt (collected near Gufunes, Reykjavik, Iceland; chosen as an analogue of Moon/Mars surface material) inside a purposely designed miniature biomining reactor housed in the KUBIK incubator. Sphingomonas desiccabilis enhanced mean leached concentrations of rare earth elements (the lanthanides plus yttrium) relative to non-biological controls in all gravity regimes, while Bacillus subtilis reduced REE bioleaching efficacy and Cupriavidus metallidurans showed no difference, demonstrating the microbial specificity of the process as on Earth. A companion analysis showed that Sphingomonas desiccabilis and Bacillus subtilis both enhanced vanadium leaching by 184.92 to 283.22% relative to sterile controls, and a third study found no significant differences in final bacterial cell concentrations between gravity regimes. Gravity level had no significant effect on mean leaching in any case, establishing the principle of biomining and associated bioindustrial processes on Solar System bodies with non-1 x g gravity.

Taxonomy

Taxon Ontology ID Functional Roles Abundance
Sphingomonas desiccabilis CP1D NCBITaxon:429134
PRIMARY_DEGRADER
N/A
  • PMID:33173035 - SUPPORT (IN_VITRO)
    "Sphingomonas desiccabilis CP1D (DSM 16792; Type strain)"
  • PMID:33173035 - SUPPORT (IN_VITRO)
    "Sphingomonas desiccabilis enhanced mean leached concentrations of REEs compared to non-biological controls in all gravity conditions"
Bacillus subtilis NCIB 3610 NCBITaxon:1423
PRIMARY_DEGRADER
N/A
  • PMID:33173035 - SUPPORT (IN_VITRO)
    "Bacillus subtilis NCIB 3610 (DSM 10; Type strain)"
  • PMID:33868198 - SUPPORT (IN_VITRO)
    "Sphingomonas desiccabilis and Bacillus subtilis enhanced the leaching of vanadium under the three gravity conditions compared to sterile controls by 184.92 to 283.22%"
Cupriavidus metallidurans CH34 NCBITaxon:119219 N/A
  • PMID:33173035 - SUPPORT (IN_VITRO)
    "Cupriavidus metallidurans CH34 (DSM 2839; Type strain)"
  • PMID:33154740 - SUPPORT (IN_VITRO)
    "Cupriavidus metallidurans CH34"

Ecological Interactions

Ecological interaction network for BioRock ISS Basalt Biomining Consortium Bipartite graph where circle nodes represent taxa and colored rectangles represent ecological interactions (cross-feeding, mutualism, syntrophy, competition, commensalism).
Taxon
Cross-feeding
Mutualism
Syntrophy
Competition
Commensalism
Niche partitioning
Colonization facilitation
Strain competition
Predation

Sphingomonas desiccabilis rare-earth-element bioleaching from basalt

CROSS_FEEDING

Source Taxon: Sphingomonas desiccabilis CP1D

Metabolites: lanthanum(3+) (CHEBI:49701), cerium(3+) (CHEBI:48782), neodymium(3+) (CHEBI:229785), yttrium(3+) (CHEBI:49962)

Biological Processes:

Evidence

  • PMID:33173035 - SUPPORT (IN_VITRO)
    "Sphingomonas desiccabilis enhanced mean leached concentrations of REEs compared to non-biological controls in all gravity conditions"
  • PMID:33173035 - SUPPORT (IN_VITRO)
    "Content of rare earth elements (REEs; reported as μg/g"

Vanadium bioleaching by Sphingomonas desiccabilis and Bacillus subtilis

CROSS_FEEDING

Source Taxon: Bacillus subtilis NCIB 3610

Biological Processes:

Evidence

  • PMID:33868198 - SUPPORT (IN_VITRO)
    "Sphingomonas desiccabilis and Bacillus subtilis enhanced the leaching of vanadium under the three gravity conditions compared to sterile controls by 184.92 to 283.22%"
  • PMID:33868198 - SUPPORT (IN_VITRO)
    "One element of interest in mining is vanadium (V)"

Microbial specificity of basalt bioleaching among panel members

NICHE_PARTITIONING

Source Taxon: Sphingomonas desiccabilis CP1D

Target Taxon: Cupriavidus metallidurans CH34

Evidence

  • PMID:33173035 - SUPPORT (IN_VITRO)
    "Bacillus subtilis exhibited a reduction in bioleaching efficacy and Cupriavidus metallidurans showed no difference compared to non-biological controls"
  • PMID:33173035 - SUPPORT (IN_VITRO)
    "showing the microbial specificity of the process, as on Earth"

External Resources

Name Repository Resource ID
Primary publication - REE extraction in microgravity and Mars gravity
Nature Communications 2020 primary publication reporting rare earth element bioleaching from basalt by the BioRock panel on the ISS.
OTHER doi:10.1038/s41467-020-19276-w
  • PMID:33173035 - SUPPORT (IN_VITRO)
    "bioleaching of REEs from basaltic rock"
Companion publication - vanadium mining and bioremediation
Frontiers in Microbiology 2021 companion publication reporting microbially-enhanced vanadium leaching by the BioRock panel under micro- and Mars gravity.
OTHER doi:10.3389/fmicb.2021.641387
  • PMID:33868198 - SUPPORT (IN_VITRO)
    "One element of interest in mining is vanadium (V)"
Companion publication - final bacterial cell concentrations
Frontiers in Microbiology 2020 companion publication reporting no significant effect of microgravity or simulated Mars gravity on final bacterial cell concentrations, with detailed cultivation methods for the three-species panel.
OTHER doi:10.3389/fmicb.2020.579156
  • PMID:33154740 - SUPPORT (IN_VITRO)
    "no significant differences in final cell counts and optical densities between the three gravity regimens on the ISS"
NCBI Taxonomy - Sphingomonas desiccabilis
NCBI Taxonomy record for Sphingomonas desiccabilis.
OTHER NCBITaxon:429134
NCBI Taxonomy - Bacillus subtilis
NCBI Taxonomy record for Bacillus subtilis.
OTHER NCBITaxon:1423
NCBI Taxonomy - Cupriavidus metallidurans
NCBI Taxonomy record for Cupriavidus metallidurans.
OTHER NCBITaxon:119219

Environmental Factors

Factor Value Unit
Gravity regime microgravity, simulated Mars gravity, and simulated Earth gravity (plus ground 1 x g controls) N/A
  • PMID:33154740 - SUPPORT (IN_VITRO)
    "microbe-mineral interactions in microgravity, simulated Mars gravity and simulated Earth gravity"
Basalt substrate olivine basalt slides (Gufunes, Reykjavik, Iceland) N/A
  • PMID:33173035 - SUPPORT (IN_VITRO)
    "olivine basalt rock collected near Gufunes, Reykjavik in Iceland"
Incubation temperature and duration 21 days at ~20 C N/A
  • PMID:33154740 - SUPPORT (IN_VITRO)
    "Samples grew for 21 days at 20.16°C"

Growth Media