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High-Solids Switchgrass Methanogenic Microbiome

A thermophilic, anaerobic, semi-continuously fed methanogenic enrichment microbiome cultivated on mid-season switchgrass at high solids loadings. The community maintained fractional carbohydrate solubilization as solids loading increased from 30 to 150 g/L, with methane and carbon dioxide as the main steady-state products. Metaproteomics identified bacterial carbohydrate-active enzyme strategies and archaeal methanogenesis functions relevant to DOE bioenergy efforts to understand robust lignocellulose deconstruction under industrially relevant process conditions.

Taxonomy

Taxon Ontology ID Functional Roles Abundance
Clostridia NCBITaxon:186801
PRIMARY_DEGRADER SECONDARY_FERMENTER
DOMINANT
  • PMID:35790765 - SUPPORT (COMPUTATIONAL)
    "Firmicutes (Clostridia followed by Bacillus)"
Bacilli NCBITaxon:91061
PRIMARY_DEGRADER SECONDARY_FERMENTER
ABUNDANT
  • PMID:35790765 - SUPPORT (COMPUTATIONAL)
    "Clostridia followed by Bacillus"
Chloroflexota NCBITaxon:200795
PRIMARY_DEGRADER SECONDARY_FERMENTER
COMMON
  • PMID:35790765 - SUPPORT (COMPUTATIONAL)
    "phylum Chloroflexi were more prevalent"
Thermotogota NCBITaxon:200918
PRIMARY_DEGRADER SECONDARY_FERMENTER
COMMON
Euryarchaeota NCBITaxon:28890
SYNTROPHIC_PARTNER CROSS_FEEDER
COMMON
  • PMID:35790765 - SUPPORT (COMPUTATIONAL)
    "Euryarchaeota proteins"

Ecological Interactions

Ecological interaction network for High-Solids Switchgrass Methanogenic Microbiome Bipartite graph where circle nodes represent taxa and colored rectangles represent ecological interactions (cross-feeding, mutualism, syntrophy, competition, commensalism).
Taxon
Cross-feeding
Mutualism
Syntrophy
Competition
Commensalism
Niche partitioning
Colonization facilitation
Strain competition
Predation

High-Solids Switchgrass Carbohydrate Solubilization

NICHE_PARTITIONING

Source Taxon: Clostridia

Metabolites: cellulose (CHEBI:18246), xylan (CHEBI:37686)

Biological Processes:

Evidence

  • PMID:35790765 - SUPPORT (IN_VITRO)
    "undiminished fractional carbohydrate solubilization"
  • PMID:35790765 - SUPPORT (COMPUTATIONAL)
    "specific carbohydrate-active enzyme classes"

Fraction-Specific CAZyme Functional Niches

NICHE_PARTITIONING

Source Taxon: Chloroflexota

Target Taxon: Thermotogota

Metabolites: cellulose (CHEBI:18246), xylan (CHEBI:37686)

Biological Processes:

  • biological process involved in interspecies interaction between organisms (GO:0044419)

Evidence

  • PMID:35790765 - SUPPORT (COMPUTATIONAL)
    "suggesting the presence of functional niches"
  • PMID:35790765 - SUPPORT (COMPUTATIONAL)
    "substrate bound"

Hydrogenotrophic Methanogenic Sink

SYNTROPHY

Source Taxon: Euryarchaeota

Metabolites: methane (CHEBI:16183), carbon dioxide (CHEBI:16526), dihydrogen (CHEBI:18276)

Biological Processes:

Evidence

  • PMID:35790765 - SUPPORT (IN_VITRO)
    "The main products observed were methane"
  • PMID:35790765 - SUPPORT (COMPUTATIONAL)
    "all hydrogenotrophic methanogenesis pathway enzymes"

Associated Datasets

Dataset Type Repository Accession
High-solids microbiome MassIVE metaproteomics dataset
Raw mass spectra for the metaproteomics study of the high-solids switchgrass microbiome.
METAPROTEOME OTHER MSV000088319
High-solids microbiome ProteomeXchange dataset
ProteomeXchange record for the metaproteomics data from the high-solids switchgrass microbiome.
METAPROTEOME OTHER PXD029582

External Resources

Name Repository Resource ID
Exact-system primary publication - high-solids switchgrass microbiome
Primary ORNL/NREL-associated publication reporting metaproteomics of the anaerobic thermophilic switchgrass microbiome at increasing solids loading.
OTHER PMID:35790765
  • PMID:35790765 - SUPPORT (IN_VITRO)
    "anaerobic methanogenic microbiome"
Nature Communications full-text article
Open-access article with methods, data availability, and supplementary metaproteomics tables.
OTHER doi:10.1038/s41467-022-31433-x
MassIVE dataset DOI
Dataset DOI for raw metaproteomics mass spectra.
OTHER doi:10.25345/C5H85T

Environmental Factors

Factor Value Unit
Switchgrass solids loading 30 to 150 g/L
Anaerobic thermophilic operation anaerobic thermophilic enrichment N/A
  • PMID:35790765 - SUPPORT (IN_VITRO)
    "anaerobic, thermophilic, semi-continuously fed"
  • PMID:35790765 - SUPPORT (IN_VITRO)
    "utilize lignocellulose at high solids loadings"
High-solids enzyme response increased AA6, beta-glucosidase, xylosidase, debranching, and pectin-acting enzyme abundance N/A
  • PMID:35790765 - SUPPORT (COMPUTATIONAL)
    "auxiliary activity family 6 enzymes"
  • PMID:35790765 - SUPPORT (COMPUTATIONAL)
    "xylosidases, carbohydrate-debranching, and pectin-acting enzymes"

Growth Media