NAD salvage pathway IV (from nicotinamide riboside)

gomodel:YeastPathways_PWY3O-4106

NAD salvage pathway IV from nicotinamide riboside in Saccharomyces cerevisiae S288C uses nicotinamide riboside kinase and nicotinamide-nucleotide adenylyltransferase activities to convert N-ribosylnicotinamide through NMN to NAD.

Pathway type

nad salvage pathway iv

Organism scope

Saccharomyces cerevisiae S288C NCBITaxon:559292

Read the source YAML

Pathway network

19 nodes · 28 asserted directed edges

Arrows show recorded subject → object relationships, not necessarily metabolic flow. Select a node for details or an arrow for evidence. Scroll, zoom or find a node to explore.

Reaction / activity
Chemical / cofactor
Protein / complex
Cell / process context
Solid arrow: mechanistic relation
Dashed arrow: membership, location or cofactor relation

Show at full size (scroll to explore)

Components and activities

Declared biological components and source reaction directions
Node	Kind or role	Source direction
Saccharomyces cerevisiae S288C `NCBITaxon:559292`	taxon
hydron `CHEBI:15378`	small molecule
cytosol `GO:0005829`	cellular component
diphosphoric acid `CHEBI:29888`	small molecule
NMN(-) `CHEBI:14649`	small molecule
NRK1 Scer `SGD:S000005073`	protein
NAD(1-) `CHEBI:57540`	small molecule
N-ribosylnicotinamide `CHEBI:15927`	small molecule
ATP(4-) `CHEBI:30616`	small molecule
NMA1 Scer `SGD:S000004320`	protein
NAD+ biosynthetic process via the salvage pathway `GO:0034355`	biological process
ADP(3-) `CHEBI:456216`	small molecule
NMA2 Scer `SGD:S000003242`	protein
cytoplasm `GO:0005737`	cellular component
nucleus `GO:0005634`	cellular component
nickel(2+) `CHEBI:49786`	cofactor
cobalt(2+) `CHEBI:48828`	cofactor
β-nicotinamide D-ribonucleotide + ATP + H⁺ -> NAD⁺ + diphosphate `gomodel:2.7.7.1-RXN`	molecular activity
1-(β-D ribofuranosyl)nicotinamide + ATP -> β-nicotinamide D-ribonucleotide + ADP + H⁺ `gomodel:RIBOSYLNICOTINAMIDE-KINASE-RXN`	molecular activity
nicotinamide-nucleotide adenylyltransferase activity `gomodel:YeastPathways_PWY3O-4106/6a46ef0600000917`	molecular activity

Mechanistic edges

Mechanistic edges and their cited evidence
Subject	Predicate and description	Object	Evidence
1-(β-D ribofuranosyl)nicotinamide + ATP -> β-nicotinamide D-ribonucleotide + ADP + H⁺ `gomodel:RIBOSYLNICOTINAMIDE-KINASE-RXN`	has_output	hydron `CHEBI:15378`	gomodel:YeastPathways_PWY3O-4106 Source assertion: The model asserts gomodel:RIBOSYLNICOTINAMIDE-KINASE-RXN RO:0002234 gomodel:CHEBI_15378_RIBOSYLNICOTINAMIDE-KINASE-RXN. Source location `./YeastPathways_PWY3O-4106.json#/facts/0; endpoint types: /individuals/10/type; /individuals/2/type`
β-nicotinamide D-ribonucleotide + ATP + H⁺ -> NAD⁺ + diphosphate `gomodel:2.7.7.1-RXN`	has_input	hydron `CHEBI:15378`	gomodel:YeastPathways_PWY3O-4106 Source assertion: The model asserts gomodel:2.7.7.1-RXN RO:0002233 gomodel:CHEBI_15378_2.7.7.1-RXN. Source location `./YeastPathways_PWY3O-4106.json#/facts/1; endpoint types: /individuals/28/type; /individuals/1/type`
β-nicotinamide D-ribonucleotide + ATP + H⁺ -> NAD⁺ + diphosphate `gomodel:2.7.7.1-RXN`	occurs_in	cytosol `GO:0005829`	gomodel:YeastPathways_PWY3O-4106 Source assertion: The model asserts gomodel:2.7.7.1-RXN BFO:0000066 gomodel:reaction_2.7.7.1-RXN_location_lociGO_0005829. Source location `./YeastPathways_PWY3O-4106.json#/facts/2; endpoint types: /individuals/28/type; /individuals/26/type`
nicotinamide-nucleotide adenylyltransferase activity `gomodel:YeastPathways_PWY3O-4106/6a46ef0600000917`	has_input	hydron `CHEBI:15378`	gomodel:YeastPathways_PWY3O-4106 Source assertion: The model asserts gomodel:YeastPathways_PWY3O-4106/6a46ef0600000917 RO:0002233 gomodel:YeastPathways_PWY3O-4106/6a46ef0600000918. Source location `./YeastPathways_PWY3O-4106.json#/facts/3; endpoint types: /individuals/30/type; /individuals/31/type`
β-nicotinamide D-ribonucleotide + ATP + H⁺ -> NAD⁺ + diphosphate `gomodel:2.7.7.1-RXN`	has_output	diphosphoric acid `CHEBI:29888`	gomodel:YeastPathways_PWY3O-4106 Source assertion: The model asserts gomodel:2.7.7.1-RXN RO:0002234 gomodel:CHEBI_29888_2.7.7.1-RXN. Source location `./YeastPathways_PWY3O-4106.json#/facts/4; endpoint types: /individuals/28/type; /individuals/4/type`
1-(β-D ribofuranosyl)nicotinamide + ATP -> β-nicotinamide D-ribonucleotide + ADP + H⁺ `gomodel:RIBOSYLNICOTINAMIDE-KINASE-RXN`	has_output	NMN(-) `CHEBI:14649`	gomodel:YeastPathways_PWY3O-4106 Source assertion: The model asserts gomodel:RIBOSYLNICOTINAMIDE-KINASE-RXN RO:0002234 gomodel:CHEBI_14649_RIBOSYLNICOTINAMIDE-KINASE-RXN. Source location `./YeastPathways_PWY3O-4106.json#/facts/5; endpoint types: /individuals/10/type; /individuals/0/type`
NRK1 Scer `SGD:S000005073`	enables	1-(β-D ribofuranosyl)nicotinamide + ATP -> β-nicotinamide D-ribonucleotide + ADP + H⁺ `gomodel:RIBOSYLNICOTINAMIDE-KINASE-RXN`	gomodel:YeastPathways_PWY3O-4106 Source assertion: The model asserts gomodel:RIBOSYLNICOTINAMIDE-KINASE-RXN RO:0002333 gomodel:MONOMER3O-4139_RIBOSYLNICOTINAMIDE-KINASE-RXN_controller. Source location `./YeastPathways_PWY3O-4106.json#/facts/6; endpoint types: /individuals/10/type; /individuals/9/type`
β-nicotinamide D-ribonucleotide + ATP + H⁺ -> NAD⁺ + diphosphate `gomodel:2.7.7.1-RXN`	has_output	NAD(1-) `CHEBI:57540`	gomodel:YeastPathways_PWY3O-4106 Source assertion: The model asserts gomodel:2.7.7.1-RXN RO:0002234 gomodel:CHEBI_57540_2.7.7.1-RXN. Source location `./YeastPathways_PWY3O-4106.json#/facts/7; endpoint types: /individuals/28/type; /individuals/8/type`
1-(β-D ribofuranosyl)nicotinamide + ATP -> β-nicotinamide D-ribonucleotide + ADP + H⁺ `gomodel:RIBOSYLNICOTINAMIDE-KINASE-RXN`	has_input	N-ribosylnicotinamide `CHEBI:15927`	gomodel:YeastPathways_PWY3O-4106 Source assertion: The model asserts gomodel:RIBOSYLNICOTINAMIDE-KINASE-RXN RO:0002233 gomodel:CHEBI_15927_RIBOSYLNICOTINAMIDE-KINASE-RXN. Source location `./YeastPathways_PWY3O-4106.json#/facts/8; endpoint types: /individuals/10/type; /individuals/3/type`
nicotinamide-nucleotide adenylyltransferase activity `gomodel:YeastPathways_PWY3O-4106/6a46ef0600000917`	has_input	ATP(4-) `CHEBI:30616`	gomodel:YeastPathways_PWY3O-4106 Source assertion: The model asserts gomodel:YeastPathways_PWY3O-4106/6a46ef0600000917 RO:0002233 gomodel:YeastPathways_PWY3O-4106/6a46ef0600000919. Source location `./YeastPathways_PWY3O-4106.json#/facts/9; endpoint types: /individuals/30/type; /individuals/32/type`
NMA1 Scer `SGD:S000004320`	enables	β-nicotinamide D-ribonucleotide + ATP + H⁺ -> NAD⁺ + diphosphate `gomodel:2.7.7.1-RXN`	gomodel:YeastPathways_PWY3O-4106 Source assertion: The model asserts gomodel:2.7.7.1-RXN RO:0002333 gomodel:YLR328W-MONOMER_2.7.7.1-RXN_controller. Source location `./YeastPathways_PWY3O-4106.json#/facts/10; endpoint types: /individuals/28/type; /individuals/11/type`
β-nicotinamide D-ribonucleotide + ATP + H⁺ -> NAD⁺ + diphosphate `gomodel:2.7.7.1-RXN`	has_input	ATP(4-) `CHEBI:30616`	gomodel:YeastPathways_PWY3O-4106 Source assertion: The model asserts gomodel:2.7.7.1-RXN RO:0002233 gomodel:CHEBI_30616_2.7.7.1-RXN. Source location `./YeastPathways_PWY3O-4106.json#/facts/11; endpoint types: /individuals/28/type; /individuals/5/type`
1-(β-D ribofuranosyl)nicotinamide + ATP -> β-nicotinamide D-ribonucleotide + ADP + H⁺ `gomodel:RIBOSYLNICOTINAMIDE-KINASE-RXN`	has_input	ATP(4-) `CHEBI:30616`	gomodel:YeastPathways_PWY3O-4106 Source assertion: The model asserts gomodel:RIBOSYLNICOTINAMIDE-KINASE-RXN RO:0002233 gomodel:CHEBI_30616_RIBOSYLNICOTINAMIDE-KINASE-RXN. Source location `./YeastPathways_PWY3O-4106.json#/facts/13; endpoint types: /individuals/10/type; /individuals/6/type`
1-(β-D ribofuranosyl)nicotinamide + ATP -> β-nicotinamide D-ribonucleotide + ADP + H⁺ `gomodel:RIBOSYLNICOTINAMIDE-KINASE-RXN`	occurs_in	cytosol `GO:0005829`	gomodel:YeastPathways_PWY3O-4106 Source assertion: The model asserts gomodel:RIBOSYLNICOTINAMIDE-KINASE-RXN BFO:0000066 gomodel:reaction_RIBOSYLNICOTINAMIDE-KINASE-RXN_location_lociGO_0005829. Source location `./YeastPathways_PWY3O-4106.json#/facts/14; endpoint types: /individuals/10/type; /individuals/27/type`
nicotinamide-nucleotide adenylyltransferase activity `gomodel:YeastPathways_PWY3O-4106/6a46ef0600000917`	has_output	NAD(1-) `CHEBI:57540`	gomodel:YeastPathways_PWY3O-4106 Source assertion: The model asserts gomodel:YeastPathways_PWY3O-4106/6a46ef0600000917 RO:0002234 gomodel:YeastPathways_PWY3O-4106/6a46ef0600000921. Source location `./YeastPathways_PWY3O-4106.json#/facts/15; endpoint types: /individuals/30/type; /individuals/34/type`
nicotinamide-nucleotide adenylyltransferase activity `gomodel:YeastPathways_PWY3O-4106/6a46ef0600000917`	has_output	diphosphoric acid `CHEBI:29888`	gomodel:YeastPathways_PWY3O-4106 Source assertion: The model asserts gomodel:YeastPathways_PWY3O-4106/6a46ef0600000917 RO:0002234 gomodel:YeastPathways_PWY3O-4106/6a46ef0600000920. Source location `./YeastPathways_PWY3O-4106.json#/facts/16; endpoint types: /individuals/30/type; /individuals/33/type`
nicotinamide-nucleotide adenylyltransferase activity `gomodel:YeastPathways_PWY3O-4106/6a46ef0600000917`	has_input	NMN(-) `CHEBI:14649`	gomodel:YeastPathways_PWY3O-4106 Source assertion: The model asserts gomodel:YeastPathways_PWY3O-4106/6a46ef0600000917 RO:0002233 gomodel:CHEBI_14649_RIBOSYLNICOTINAMIDE-KINASE-RXN. Source location `./YeastPathways_PWY3O-4106.json#/facts/17; endpoint types: /individuals/30/type; /individuals/0/type`
β-nicotinamide D-ribonucleotide + ATP + H⁺ -> NAD⁺ + diphosphate `gomodel:2.7.7.1-RXN`	part_of	NAD+ biosynthetic process via the salvage pathway `GO:0034355`	gomodel:YeastPathways_PWY3O-4106 Source assertion: The model asserts gomodel:2.7.7.1-RXN BFO:0000050 gomodel:YeastPathways_PWY3O-4106/YeastPathways_PWY3O-4106. Source location `./YeastPathways_PWY3O-4106.json#/facts/18; endpoint types: /individuals/28/type; /individuals/29/type`
1-(β-D ribofuranosyl)nicotinamide + ATP -> β-nicotinamide D-ribonucleotide + ADP + H⁺ `gomodel:RIBOSYLNICOTINAMIDE-KINASE-RXN`	has_output	ADP(3-) `CHEBI:456216`	gomodel:YeastPathways_PWY3O-4106 Source assertion: The model asserts gomodel:RIBOSYLNICOTINAMIDE-KINASE-RXN RO:0002234 gomodel:CHEBI_456216_RIBOSYLNICOTINAMIDE-KINASE-RXN. Source location `./YeastPathways_PWY3O-4106.json#/facts/19; endpoint types: /individuals/10/type; /individuals/7/type`
1-(β-D ribofuranosyl)nicotinamide + ATP -> β-nicotinamide D-ribonucleotide + ADP + H⁺ `gomodel:RIBOSYLNICOTINAMIDE-KINASE-RXN`	part_of	NAD+ biosynthetic process via the salvage pathway `GO:0034355`	gomodel:YeastPathways_PWY3O-4106 Source assertion: The model asserts gomodel:RIBOSYLNICOTINAMIDE-KINASE-RXN BFO:0000050 gomodel:YeastPathways_PWY3O-4106/YeastPathways_PWY3O-4106. Source location `./YeastPathways_PWY3O-4106.json#/facts/20; endpoint types: /individuals/10/type; /individuals/29/type`
NMA2 Scer `SGD:S000003242`	enables	nicotinamide-nucleotide adenylyltransferase activity `gomodel:YeastPathways_PWY3O-4106/6a46ef0600000917`	gomodel:YeastPathways_PWY3O-4106 Source assertion: The model asserts gomodel:YeastPathways_PWY3O-4106/6a46ef0600000917 RO:0002333 gomodel:YeastPathways_PWY3O-4106/6a46ef0600000923. Source location `./YeastPathways_PWY3O-4106.json#/facts/21; endpoint types: /individuals/30/type; /individuals/35/type`
β-nicotinamide D-ribonucleotide + ATP + H⁺ -> NAD⁺ + diphosphate `gomodel:2.7.7.1-RXN`	has_input	NMN(-) `CHEBI:14649`	gomodel:YeastPathways_PWY3O-4106 Source assertion: The model asserts gomodel:2.7.7.1-RXN RO:0002233 gomodel:CHEBI_14649_RIBOSYLNICOTINAMIDE-KINASE-RXN. Source location `./YeastPathways_PWY3O-4106.json#/facts/22; endpoint types: /individuals/28/type; /individuals/0/type`
nicotinamide-nucleotide adenylyltransferase activity `gomodel:YeastPathways_PWY3O-4106/6a46ef0600000917`	part_of	NAD+ biosynthetic process via the salvage pathway `GO:0034355`	gomodel:YeastPathways_PWY3O-4106 Source assertion: The model asserts gomodel:YeastPathways_PWY3O-4106/6a46ef0600000917 BFO:0000050 gomodel:YeastPathways_PWY3O-4106/YeastPathways_PWY3O-4106. Source location `./YeastPathways_PWY3O-4106.json#/facts/23; endpoint types: /individuals/30/type; /individuals/29/type`
NMA1 Scer `SGD:S000004320`	located_in Physical protein location from reviewed UniProt. ECO:0000269 (PubMed:14562095)	cytoplasm `GO:0005737`	UniProtKB:Q06178 Source assertion: UniProtKB:Q06178 annotates location Cytoplasm (SL-0086). Source location `https://rest.uniprot.org/uniprotkb/stream?query=taxonomy_id%3A559292+AND+reviewed%3Atrue&format=json&fields=accession%2Cprotein_name%2Cxref_sgd%2Ccc_subcellular_location%2Ccc_function%2Ccc_caution%2Corganism_id#/results/2726/comments/1/subcellularLocations/0; sha256=95f72dcc644358c480e0e59f996cf88533558b5fa4256460a030e0a1eaf9678d` UniProtKB:Q06178 Source assertion: UniProtKB:Q06178 has the exact SGD cross-reference SGD:S000004320. Source location `https://rest.uniprot.org/uniprotkb/stream?query=taxonomy_id%3A559292+AND+reviewed%3Atrue&format=json&fields=accession%2Cprotein_name%2Cxref_sgd%2Ccc_subcellular_location%2Ccc_function%2Ccc_caution%2Corganism_id#/results/2726/uniProtKBCrossReferences/0; sha256=95f72dcc644358c480e0e59f996cf88533558b5fa4256460a030e0a1eaf9678d`
NMA1 Scer `SGD:S000004320`	located_in Physical protein location from reviewed UniProt. ECO:0000269 (PubMed:14562095)	nucleus `GO:0005634`	UniProtKB:Q06178 Source assertion: UniProtKB:Q06178 annotates location Nucleus (SL-0191). Source location `https://rest.uniprot.org/uniprotkb/stream?query=taxonomy_id%3A559292+AND+reviewed%3Atrue&format=json&fields=accession%2Cprotein_name%2Cxref_sgd%2Ccc_subcellular_location%2Ccc_function%2Ccc_caution%2Corganism_id#/results/2726/comments/1/subcellularLocations/1; sha256=95f72dcc644358c480e0e59f996cf88533558b5fa4256460a030e0a1eaf9678d` UniProtKB:Q06178 Source assertion: UniProtKB:Q06178 has the exact SGD cross-reference SGD:S000004320. Source location `https://rest.uniprot.org/uniprotkb/stream?query=taxonomy_id%3A559292+AND+reviewed%3Atrue&format=json&fields=accession%2Cprotein_name%2Cxref_sgd%2Ccc_subcellular_location%2Ccc_function%2Ccc_caution%2Corganism_id#/results/2726/uniProtKBCrossReferences/0; sha256=95f72dcc644358c480e0e59f996cf88533558b5fa4256460a030e0a1eaf9678d`
NMA2 Scer `SGD:S000003242`	located_in Physical protein location from reviewed UniProt. ECO:0000269 (PubMed:11884393)	nucleus `GO:0005634`	UniProtKB:P53204 Source assertion: UniProtKB:P53204 annotates location Nucleus (SL-0191). Source location `https://rest.uniprot.org/uniprotkb/stream?query=taxonomy_id%3A559292+AND+reviewed%3Atrue&format=json&fields=accession%2Cprotein_name%2Cxref_sgd%2Ccc_subcellular_location%2Ccc_function%2Ccc_caution%2Corganism_id#/results/2159/comments/1/subcellularLocations/0; sha256=95f72dcc644358c480e0e59f996cf88533558b5fa4256460a030e0a1eaf9678d` UniProtKB:P53204 Source assertion: UniProtKB:P53204 has the exact SGD cross-reference SGD:S000003242. Source location `https://rest.uniprot.org/uniprotkb/stream?query=taxonomy_id%3A559292+AND+reviewed%3Atrue&format=json&fields=accession%2Cprotein_name%2Cxref_sgd%2Ccc_subcellular_location%2Ccc_function%2Ccc_caution%2Corganism_id#/results/2159/uniProtKBCrossReferences/0; sha256=95f72dcc644358c480e0e59f996cf88533558b5fa4256460a030e0a1eaf9678d`
NMA1 Scer `SGD:S000004320`	has_cofactor UniProtKB reviewed (Swiss-Prot) annotation. UniProt evidence: ECO:0000269 (PubMed:10428462) Divalent metal cation. UniProt note evidence: ECO:0000269 (PubMed:10428462)	nickel(2+) `CHEBI:49786`	UniProtKB:Q06178 Source assertion: UniProtKB:Q06178 annotates CHEBI:49786 as a cofactor of this protein. Source location `https://rest.uniprot.org/uniprotkb/stream?query=taxonomy_id%3A559292+AND+reviewed%3Atrue&format=json&fields=accession%2Cprotein_name%2Cxref_sgd%2Ccc_cofactor%2Corganism_id%2Ccc_caution [sha256:1378282d22c5518215707964664b55946f468ba74a227a6ea1bbf27db69ddbc4] #/results/2726/comments/0/cofactors/0; /results/2726/comments/0/note` UniProtKB:Q06178 Source assertion: UniProtKB:Q06178 has the exact SGD cross-reference SGD:S000004320. Source location `https://rest.uniprot.org/uniprotkb/stream?query=taxonomy_id%3A559292+AND+reviewed%3Atrue&format=json&fields=accession%2Cprotein_name%2Cxref_sgd%2Ccc_cofactor%2Corganism_id%2Ccc_caution [sha256:1378282d22c5518215707964664b55946f468ba74a227a6ea1bbf27db69ddbc4] #/results/2726/uniProtKBCrossReferences/0`
NMA2 Scer `SGD:S000003242`	has_cofactor UniProtKB reviewed (Swiss-Prot) annotation. UniProt evidence: ECO:0000269 (PubMed:12597897) Divalent metal cation. UniProt note evidence: ECO:0000269 (PubMed:12597897)	cobalt(2+) `CHEBI:48828`	UniProtKB:P53204 Source assertion: UniProtKB:P53204 annotates CHEBI:48828 as a cofactor of this protein. Source location `https://rest.uniprot.org/uniprotkb/stream?query=taxonomy_id%3A559292+AND+reviewed%3Atrue&format=json&fields=accession%2Cprotein_name%2Cxref_sgd%2Ccc_cofactor%2Corganism_id%2Ccc_caution [sha256:1378282d22c5518215707964664b55946f468ba74a227a6ea1bbf27db69ddbc4] #/results/2159/comments/0/cofactors/0; /results/2159/comments/0/note` UniProtKB:P53204 Source assertion: UniProtKB:P53204 has the exact SGD cross-reference SGD:S000003242. Source location `https://rest.uniprot.org/uniprotkb/stream?query=taxonomy_id%3A559292+AND+reviewed%3Atrue&format=json&fields=accession%2Cprotein_name%2Cxref_sgd%2Ccc_cofactor%2Corganism_id%2Ccc_caution [sha256:1378282d22c5518215707964664b55946f468ba74a227a6ea1bbf27db69ddbc4] #/results/2159/uniProtKBCrossReferences/0`

References

gomodel:YeastPathways_PWY3O-4106 SGD GO-CAM NAD salvage pathway IV (from nicotinamide riboside) - imported from: Saccharomyces Genome Database (source version: Pinned Noctua JSON archive; model date 2026-07-06; member ./YeastPathways_PWY3O-4106.json)
UniProtKB:Q06178 Reviewed UniProtKB Q06178 location annotations (source version: UniProt release 2026_03)
UniProtKB:P53204 Reviewed UniProtKB P53204 location annotations (source version: UniProt release 2026_03)

Curation history

2026-10-05T06:05:08Z Codex; review-causal-graph; Reviewed every native GO-CAM fact; preserved precise input/output, causal, location, membership and complex-part relations with traceable structured source evidence. Kept distinct generic physical instances and recorded source-scope exclusions in the cohort review ledger.
2026-10-05T06:43:56Z Codex
- EDIT — Audit causal activity compartments and preserve scoped protein location evidence
  Reviewed all 3 activity location assignments and 3 mapped physical proteins against reviewed S288C UniProt release2026_03 location/function/caution batch SHA256 95f72dcc644358c480e0e59f996cf88533558b5fa4256460a030e0a1eaf9678d. Added 3 independently supported location edges and quarantined 1 generic cytosol source assertions. Mapped physical protein locations via exact SGD cross-references; retained source ECO grades, molecular form and conditional notes. Physical protein location alone never generates activity location. Explicitly inspected functional assertions support bounded activity projections. Every original activity location and exclusion is recorded in reports/causal_graph_review/gocam-location-review.json. No independent location annotation is treated as an evidence gap, not a negative result. Cytoplasm is not narrowed to cytosol. Microsome fractions are not projected as in-vivo organelles. Semantic validation passed all85 records;4 independent source-role acceptance tests passed.
Read this curation session